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^Ilk  Missouri 
^  Botanical 
Garden 


3  1753  00355  9298 


A  Journal  for  Botanical  Nomenclature 


2006 


Volume  16,  Number  1 
April  2006 


Novon,  A  Journal  for  Botanical 
Nomenclature  from  the  Missouri 
Botanical  Garden 


The  mission  of  the  Missouri  Botanical  Garden  is  to  discover  and  share  knowledge 
about  plants  and  their  environment,  in  order  to  preserve  and  enrich  life. 


Diana  Gunter 


Editor 


Kanchi  N.  Gandhi 


Volume  16  NO  VON  ^(|(|/ 

Number  1  ,J,ssQU»''°,fWC,L  K2& 

2006  wo.* 

GARDEN  LIBRARY - 

Inclusion  of  Suua  in  Sauvagesia  (Ochnaceae) 


No  von  16:  1-2.  Published  on  25  May  2006. 


Validation  of  Piper  methysticum  var.  wichmannii  (Piperaceae) 


The  Esparto  Grass  Question:  A  Systematic  Approach  for  a  Long-Lasting 
Problem  in  Stipa  L.  (Gramineae) 


Central  Brazil 


$%■*■?  p 


21 


A  New  Name  in  Primula  (Primulaceae) 


Komarov  Botanical  Institute,  Russian  AcatW^Sde, 


Novon 


New  Taxa  of  Anthurium  (Araceae)  from  the  Bajo  Calima  Region 
(Valle,  Choco),  Colombia  and  Ecuador 

Thomas  B.  Croat 

Missouri  Botanical  Garden,  P.0.  Box  299,  St.  Louis,  Missouri  63166-0299. 


as 


Road, 


Emily  D.  Yates 

P.O.  Box  299.  St.  Louis,  Missouri  63166-0299. 
Las  Vegas,  NV  89124.  emilydrewyates@hotmail 


i  lifilutil 


27 


32 


Ifllllli 


(T-wf/ 


i),  a  New  Species  from  Chile 


54 


Nt 


in  Erica  (Ericaceae:  Ericoideae)  from  the  High 
Mountains  of  East  Africa 


n  of  Natural  History, 

DC  20013-7012,  U.S.A. 


A  New  Species  of  Cassipourea  (Rhizophoraceae)  from 


Western  Cameroon 


K  I  UIH'iHKirKiiHHl 


A  New  Species  and  Notes  on  Drymonia  (Gesneriaceae)  from  Costa  Rica 


Ricardo  Kriebel 


^|rAn^er^ere.i withd^lal  gknd . -G.  FruiL—H.  Cross  section  ,  I  II,  Ty  endocarp.  (  \  <  I  ,  A.  Estrada  399 ; 


Validation  of  Names  at  Subspecific  Rank  in  North 
American  Campanulaceae 


16:  69-73 i  Published  on  25  May  2006. 


71 


73 


A  New  Species  of  Mimosa  (Fabaceae,  Mimosoideae)  from  Madagascar 


Volume  16,  Number  1 
2006 


Lefevre  &  Labat 

Mimosa  manomboensis  from  Madagascar 


(K  nun  ui(l<| 


■  from  Madagas 


A  New  Species  of  Caesalpinia  (Leguminosae,  Caesalpinioideae) 
from  China 


Volume  16,  Number  1 
2006 


Li  et  al. 

Caesalpinia  yunnaner 


(Rubiaceae,  Sp 
the  Mixed  Te 


e),  a  New  Species  from 
of  Mexico 


l  63,  91000  Xalapa,  Ve 


Passiflora  sandrae  (Passifloraceae),  a  New  Species  from  Panama 


Novon 


00100, 


David  A.  Simpson 


Neillia  Includes  Stephanandra  (Rosaceae) 


94 


Oh 


1 theca  (Compositae:  Astereae),  a 
Combination  for  a  Common  Neotropical  Weed 


Gisela  Sancho 


Palmorchis 


a  New  Species  of  Orohidaceae  from 


a  New  Species  < 
from  the  Sierra  Madre  Oriental,  Nuevo  Leon,  Mexico 


106 


Novon 


Novon 


x  !!■!  till 


Salcedo  et  al. 

Phaseolus  novoleonensis 


110 


Mar  Ire  Or 


mm 


et  al. 


Eight  New  Species  of  Mandevilla  Lindley  (Apocynaceae,  Apocynoideae) 
from  Brazil 


116 


Novon 


117 


Novon 


Volume  16,  Number  1 
2006 


Sales  et  al. 

Mandevilla  from  Brazil 


119 


122 


Novon 


Volume  16,  Number  1 
2006 


Sales  et  al. 

Mandevilla  from  Brazil 


CFCR  1195,  SPF,  UKC.) 


Volume  16,  Number 
2006 


ft  ft 


Guianodendron ,  a  New  Genus  of  Leguminosae  (Papilionoideae) 


130 


Novon 


(Fig.  2). 


Magnolia  amabilis ,  a  New  Species  of  Magnoliaceae  from  Yi 


China 


Sima  Yong-Kang 


State  Forestry  Administration,  Kunming  650204,  China 


Yu-Min* 


650204,  China. 


Novon 


A  New  Name  and  Three  New  Combinath 


ons  in  Brazilian  Myrtaceae 


)  Carlos  6627,  31270-110,  llelo  I 


16:  136  137.  I  ed  on  25  May  2006. 


Arnica  lanceolata  subsp.  prima,  the  Correct  Name  for  Arnica  lanceolata 
Nuttall  subsp.  amplexicaulis  (Nuttall)  Gruezo  (Asteraceae,  Heliantheae) 


140 


et  al. 


Rubiacearum  Americanarum  Magna  Hama  Pars  XVIII: 

New  Species  of  Psychotria  Subg.  Psychotria  from  Central  America 
and  Western  South  America 


5  cm 


2006 


Taylor 

Psychotria  Subg.  Psychotria 


145 


'  Subg.  Psychotria 


A  New  Species  of  Eleocharis  R.  Brown  (Cyperaceae)  from 
Southern  Brazil 


■  (!o  Sul,  91501-970,  I 


A  New  Species  of  the  Genus  Rhamnus  (R1 


?)  from  China 


Yi-Zhi  , 

r  of  Life  Science,  Innei 


( lilcil 


25  M  w 


www.mbgpress.org 


CONTENTS 


^JJjk  Missouri 
Botanical 
Garden 


A  Journal  for  Botanical  Nomenclature 


2006 


Volume  16,  Number  2 
July  2006 


Novon,  A  Journal  for  Botanical 
Nomenclature  from  the  Missouri 
Botanical  Garden 


The  mission  of  the  Missouri  Botanical  Garden  is  to  discover  and  share  knowledge 
about  plants  and  their  environment  in  order  to  preserve  and  enrich  life. 


Novon  publishes  short  articles  whose  primary  purpose  is  the  establishment  of  nomencla¬ 
ture  in  vascular  plants  and  bryophytes.  All  articles  are  peer-reviewed  by  qualified,  indepen¬ 
dent  reviewers. 

Manuscripts  must  fully  state  and  justify  the  reasons  for  proposing  nova.  These  may  include 
detailed  comparisons  with  similar  taxa,  short  keys  to  similar  taxa,  illustrations  to  similar  taxa, 
and  mechanical  nomenclature  reasons,  among  others.  Manuscripts  whose  primary  purpose  is 
other  than  establishment  of  new  nomenclature,  which  usually  are  longer  manuscripts,  cannot 
be  accepted  for  review.  These  include  reviews,  revisions,  monographs,  or  other  papers  that 
incidentally  include  nova. 

Manuscripts  must  follow  the  guidelines  in  the  Checklist  for  Authors  and  be  accompanied  by 
a  completed  copy  of  the  Checklist.  The  Checklist  may  be  downloaded  from  the  Garden’s  web 
site,  www.mbgpress.org,  or  authors  may  contact  the  managing  editor  to  request  a  copy. 

Novon  will  not  knowingly  accept  manuscripts  that  have  been  simultaneously  submitted  to 
other  journals  for  consideration  or  previously  published  in  some  form  elsewhere. 


Victoria  C.  Hollowell 

Editor 

Beth  Parada 

Managing  Editor 

Diana  Gunter 

Associate  Editor 

Ihsan  A.  Al-Shehbaz 

Consulting  Editor 

Roy  E.  Gereau 

Consulting  Editor 

John  Pruski 

Consulting  Editor 

Carmen  Ulloa  Ulloa 

Consulting  Editor 

George  Yatskievych 

Consulting  Editor 

Kanchi  N.  Gandhi 

Nomenclature  Consultant 

Nicholas  J.  Turland 

Nomenclature  Consultant 

Barbara  Mack 

Editorial  Assistant 

Monica  Anderson 

MBG  Press  Assistant 

Novon  is  included  in  the  subscription  price  of  the 
Annals  of  the  Missouri  Botanical  Garden,  Price  for 
2006:  $160  per  year  EEA.;  $170  Canada  and 
Mexico;  $195  all  other  countries.  Four  issues  per 


rence,  KS  66044-8897;  Manuscript  submissions: 
Novon,  Missouri  Botanical  Garden,  P.0.  Box  299, 
St.  Louis,  M0  63166-0299. 

Electronic  addresses: 

Editorial  queries:  novon@mobot.org 
Orders  for  back  issues:  mbgpress@mobot.org 
Web  site:  http://www.mbgpress.org 

©  Missouri  Botanical  Garden  2006 

Typesetting  by  Allen  Press,  Inc,  using  Advenl3B2  ai 


©  Novon  is  printed  on,  paper  that  meets  the  re¬ 
quirements  of  ANSI/NISO  Z39.48-1992. 

Novon  (ISSN  1055-3177)  is  published  quarterly 
by  the  Missouri  Botanical  Garden,  2345  Tower 
Grove  Avenue,  St.  Louis,  MO  63110.  Periodicals 
postage  paid  at  St,  Louis,  MO  and  additional  mail¬ 
ing  offices  (USFS  #006-777). 

POSTMASTER:  Please  send  address  changes  to 
Annals  of  the  Missouri  Botanical  Garden,  P.0.  Box 
1897,  Lawrence,  KS  66044-8897. 

Information  on  the  contents  of  Novon  is  present¬ 
ed  in  SciSearch®,  Research  Alert®,  Current  Con¬ 
tents®/ Agriculture,  Biology,  and  Environmental 
Sciences,  APT  Online,  the  CAB  Abstract/Global 
Health  database,  and  the  ISI®  database. 

'mafic  paging  system. 


Volume  16 
Number  2 
2006 


NOVON 


Solarium  perlongistylum  and  S.  catilliflorum ,  New  Endemic  Peruvian 
Species  of  Solarium ,  Section  Basarthrum ,  Are  Close  Relatives  of  the 
Domesticated  Pepino,  S.  muricatum 


Volume  16,  Number  2 


Anderson  et  al. 

Endemic  Peruvian  Species 


163 


Eight  Segregates  from  the  Neotropical  Genus  Mascagnia 
(Malpighiaceae) 


170 


171 


174 


Volume  16,  Number  2 


182 


Novon 


Novon 


12331  (MICH). 


Novon 


ill 


Volume  16,  Number  2 
2006 


Anderson 

Segregates  from  Mascagnia 


195 


Volume  16, 
2006 


201 


Segregates  from  Mascagnia 


Hits 


2006 


New  Combinations  in  Manekia ,  an  Earlier  Name  for 


Una  Nueva  Especie  de  Schwenckia  (Solanaceae)  del  Caqueta,  Colombia 

Facultad  de  Agronom.a,  Institute  de  Botanica  Agrrcola,  Universidad  Central  de  Venezuela, 
Maracay  2101  Estado  Aragua,  Venezuela.  cbenitez@telcel.net.ve 


210 


Novon 


211 


R.  G. 


Three  New  Species  of  Actinocephalus  Sano  (Eriocaulaceae)  from  Minas 
Gerais,  Brazil 


Volume  16,  Number  2 
2006 


Costa 

Actinocephalus  ■ 


213 


1.5  mm  long; 


Oresbia ,  a  New  South  African  Genus  of  the  Ast 


Volume  16,  Number  2 
2006 


Cron  &  Nordenstam 
Oresbia  (Asteraceae) 


217 


218 


Novon 


Cron  &  Nordenstam 
Oresbia  (Asteraceae) 


219 


Volume  16,  Number  2  Cron  &  Nordenstam  223 

2006  Oresbia  (Asteraceae) 


Bolandia  (As 


Eric  Knox 


If? 


Volume  16,  Number  2 
2006 


Cron  et  al. 


227 


229 


A  New  Heterophyllous  Spermacoce  Species  (Rubiaceae)  from 
the  Marungu  Highlands,  Democratic  Republic  of  the  Congo 

Steven  Dessein  and  Elmar  Robbrecht 

National  Botanic  Garden  of  Belgium,  Domein  van  Bouchout,  B-1860  Meise,  Belgium. 
steven.dessein@br.fgov.be;  elmar.robbrecht@br.fgov.be 

Erik  Smets 

Laboratory  of  Plant  Systematics,  Katholieke  Universiteit  Leuven,  Kasteelpark  Arenberg  31, 
B-3001  Leuven,  Belgium,  erik.smets@bio.kuleuven.be 


Abstract.  Spermacoce  heteromorpha  Dessein  (Ru- 
hiaceac.  Spermacoceae)  is  newly  described  and 
illustrated.  It  grows  on  the  Marungu  highlands  of  the 
Democratic  Republic  of  the  Congo  (D.R.  Congo)  at 
altitudes  of  1800  to  1900  m.  The  species  resembles 
the  Tanzanian  Spermacoce  taylorii  Verdcourt,  from 
which  it  mainly  differs  in  having  terminal  inflores¬ 
cences,  only  slightly  exserted  anthers,  and  densely 

features  marked  heterophylly,  with  long  leaves  at  ihe 

Rubiaceae  as  a  whole.  The  finding  of  this  species 
exemplifies  the  unique  floristic  richness  of  this  under¬ 
explored  part  of  D.R.  Congo. 

Key  words:  Democratic  Republic  of  the  Congo, 
heterophylly,  Marungu  highlands,  Rubiaceae,  Sper- 


Spermacoce  L.  is  a  pantropical  genus  of  the  family 

(Dessein  et  al.,  2002;  Harwood  &  Dessein,  2005). 
Tropical  America,  tropical  Africa, 
part  of  Australia  are  the  main  centers  of  diversity.  The 
genus  is  characterized  by  its  herbaceous  growth  form, 
the  fimbriate  stipules  connected  to  the  petioles,  the  4- 

a  single  ovule  per  locule,  the  often  pluriaperturate 
pollen  grains,  and  the  dry  dehiscent  fruits.  Many 

of  ruderal  places.  Typical  examples  are  S.  latifolia 

native  to  America,  but  introduced  in  Africa,  Asia,  and 
Australia.  Narrow  endemics,  however,  are  an  element 

Our  study  of  Spermacoce  from  the  high  plateaus  of  the 
Zambezian  regional  center  of  endemism,  for  example, 
revealed  the  existence  of  endemic  or  disjunct 
Spermacoce  species  on  the  Angolan,  Zambian,  and 


ly  put  at  our  disposal, 
;  from  the  Marungu 
found  representing 


Heterophylly 

Heterophylly  is  not  uncommon  among  woody 
Rubiaceae  (Robbrecht,  1988),  but  is  rarely  observed 
among  the  herbaceous  representatives  of  the  family. 
Within  Spermacoce ,  it  has  only  been  recorded  for  S. 
hockii  (De  Wildeman)  Dessein,  a  perennial  from  D.R. 
Congo  (Dessein  et  al.,  2003).  In  this  species,  long 

the  caul  me  leaves  are  broader  and  much  shorter. 
During  field  work  in  Zambia,  we  observed  that  several 
biannual  or  short-lived  perennial  Spermacoce  species 
develop  rosettes  of  basal  leaves.  This  feature  was 
observed  in  the  field  for  5.  dibrachiata  Oliver,  .S’. 


the  basal  part  is  seldom  collected.  I 
mentioned  species,  basal  rosette  leaves  differ  only 
slightly  in  size  and  shape  from  the  cauline  leaves.  In 
S.  heteromorpha,  on  the  contrary,  the  basal  leaves  are 
markedly  longer  than  the  cauline  ones.  The  type 
specimens  of  S.  taylorii  Verdcourt  also  have  basal  and 
cauline  leaves,  but  the  differences  between  the  two 
leaf  types  are  minor. 


The  Marungu  high  plateau  is  situated  southwest  of 
Lake  Tanganyika  in  D.R.  Congo.  The  name  of  the 
plateau  is  derived  from  the  word  “dilungu,”  which 
signifies  “herbaceous  plateau”  (Verbeken,  1954).  The 
name  refers  to  a  main  feature  of  the  environment,  the 
absence  of  tree  growth  in  the  more  elevated  areas. 


Novon  16:  231-234.  Published  on  26  July  2006. 


Oxypetalum  gyrophyllum  and  0.  oblanceolatum ,  New  Species  of 
Asclepiadoideae  (Apocynaceae)  from  Brazil,  and  a  Key  for 
the  0.  insigne  Group 


rap 


China 


Li  Heng  and  Dao  Zhi-Ling 


16:  240-243.  Published  on  26  July  2006. 


Li  &  Dao 


Volume  16,  Number  2 
2006 


Li  &  Dao 

Amorphophallus  xiei 


243 


Acknowledgments.  We  thank  the  reviewers  for 
providing  useful  remarks  and  suggestions.  Wilbert 
Hetterscheid  (Netherlands)  is  also  thanked  for  his 
comments  on  the  manuscript.  We  are  grateful  for  the 
assistance  and  cooperation  of  Li  Rong,  Gao  Fu,  and 


Jiang  i 


Institute 


(KIB)  in  t he  field,  Yin  Xin-Xiang  of  the  Lo 
County  Agriculture  Department  for  discussion 
the  red  konjac,  and  Ji  Yun-Heng  (KIB)  for  his  V 
preparing  this  manuscript.  The  present  stud 
sponsored  by  the  National  Science  Foun 


(U.S.A.)  grant  DKB-0103795,  and  the  Kunming 
Institute  of  Botany,  the  Chinese  Academy  of  Sciences. 
We  also  thank  Wu  Xi-Lin  (KIB)  for  preparing  the 
illustration. 


i)  from  Vietnam 


A  New  Species  of  Camellia  Section  Piquetia  (Th< 


Volume  16,  Number  2 


New  Species  of  Cybianthus  Martins  (Myrsinaceae)  from  the 
Hylaean/Andean  Interface  of  Peru 


250 


Novon 


Novon 


Brodiaea  sierrae  (Themidaceae),  a  New  Species  from 
the  Sierra  Nevada  Foothills  of  California,  U.S.A. 


Volume  16,  Number  2  Preston  255 


staminodes.  — I).  Stamen  (adaxial  view).  — E.  Stamen  (abaxial  view).  — F.  Staminode.  — G.  Pistil.  Scale  bars  equal  10  mm. 
Drawn  from  fresh  material  ( Preston  1815  [DAV]). 


42°  N 


A  New  Species  of  Manglietia  (Magnoliaceae)  from  China 


Qin  Xin-Sheng* ,  Zhou 


Xing  Fu-Wu* 


El  Genero  Gonocalyx  (Ericaceae)  en  Costa  Rica,  con  una  Nueva  Especie 


Costa  Rica.  arodrig@il 


Volume  16,  Number  2 
2006 


Rodriguez  &  Morales  265 

Gonocalyx  en  Costa  Rica 


A  New  Combination  in  Erythrina  (Leguminosae) 

Ren  Sa 

State  Key  Laboratory  of  Systematic  and  Evolutionary  Botany,  Institute  of  Botany, 
the  Chinese  Academy  of  Sciences,  Beijing  100093,  People’s  Republic  of  China. 
saren@ibcas.ac.cn 


Abstract.  A  study  of  morphological  characters 
based  on  a  review  of  herbarium  specimens  from 
Yunnan  Province,  China,  reveals  that  the  following 
new  combination  is  necessary.  The  previously  recog¬ 
nized  species  Erythrina  yunnanensis  S.  K.  Lee 
(Leguminosae)  is  reduced  in  rank  to  variety  as 
Erythrina  stricta  Roxburgh  var.  yunnanensis  (S.  K. 
Lee)  R.  Sa. 

Key  words  China,  Erythrina,  Leguminosae,  new 


Erythrina  L.  (Leguminosae)  for  an  upcoming  volume 
of  Flora  of  China,  it  was  noticed  that  the  description 
of  E.  yunnanensis  S.  K.  Lee  is  incomplete  and  the 

species  was  described  by  Lee  (1993)  from  specimens 
from  Yunnan  Province,  China.  These  collections  were 
similar  to  E.  stricta  Roxburgh  in  flower  morphology, 
especially  with  respect  to  the  spathaceous  calyx.  The 
specimens  resembled  E.  arborescens  Roxburgh  in  their 
larger  leaflets  [E.  yunnanensis  17-24.5  X  17- 
24.5  cm  and  E.  arborescens  8-25  X  8-25  cm  vs.  E. 
stricta  7-15  X  7-15  cm),  as  well  as  in  the  sparser 
distribution  of  prickles  on  branches.  However,  the  pod 
character  of  E.  yunnanensis  was  not  described  (Lee, 
1993;  Lee  &  Chang,  1995). 

author  fortunately  found  fruiting  specimens  of  E. 
yunnanensis,  which  strongly  resembled  E.  stricta.  Both 
E.  stricta  and  E.  yunnanensis  closely  resemble  each 
other  in  flower  and  pod  morphology.  The  two  taxa 
share  a  spathaceous  calyx  and  similar  pod  sizes,  with 
one  to  three  seeds  per  pod  (Table  1). 


liable  i 


Erythr 


ihology  were  much 
more  stable  as  reliable  characters  and  do  discriminate 

retain  E.  yunnanensis  at  the  specific  rank.  Its  flowers 
and  pods  are  not  distinguishable  from  E.  stricta 
(Table  1).  With  its  morphological  similarity  for 
flowers  and  pods  to  E.  stricta 
recognized  at  the  varietal  rank  v 


Erythrina  stricta  var.  yunnanensis  (S.  K.  Lee)  R. 
Sa,  comb,  et  stat.  nov.  Basionym:  Erythrina 

Guihaia  13:  101.  1993.  TYPE:  China.  Yunnan: 
Pingbian,  on  open  slope,  1400  m,  29  June  1934, 
H.  T.  Tsai  60464  (holotype,  PE;  isotypes,  PE[2]). 
Lee’s  1993  description  of  E.  stricta  var.  yunnanen- 
and  Wang  C.  W.  72059. 

leaflet  widely  remform.  oblate,  17-24.5  X  17- 
24.5  cm,  apex  with  short  caudate  10-15  mm  tip,  base 
±  cuneate;  lateral  nerves  7  to  8  on  each  side  and 
convex;  petiolule  11-15  cm  long,  near  top  dark  brown 
after  dry;  lateral  leaflet  oblate,  apex  briefly  caudate, 
base  subtruncate  to  widely  cuneate;  petiolule  1.6- 

a  raceme  with  3  to  4  flowers  clustered  on  a  common 
peduncle;  flowers  red;  peduncle  ca.  5  mm  long,  with 
dense,  dark  brown,  short  indumentum;  calyx  spatha- 


Novon  16:  267-208.  Publish 


26  July  2006. 


m  a 


268 


A  New  Species  of  Begonia  Section  Platycentrum  (Begoniaceae) 


Shui  Yu-Min 


270 


Novon 


271 


A  New  Species  of  Croton  (Euphorbiaceae)  from  Ecuador 


Volume  16,  Number  2 
2006 


Smith 


Ecuador 


273 


Croton  floccosus  from 


;  2.5-15  cm  long  with  « 


2. 5-4.1  X  1.5-2. 1  mm, 


Memecylon  batekeanum ,  a  New  Species  from  Southeastern  Gabon, 
and  a  Note  on  the  Circumscription  of  Memecylon  Sect. 
Mouririoidea  (Melastomataceae) 


P.0.  Box  299,  St.  Louis,  Missouri  63166-0299,  tJ.S.A.  gretcl 


277 


278 


Phainantha  shuariorum  (Melastomataceae),  una  Especie  Nueva  de  la 
Cordillera  del  Condor,  Ecuador,  Disyunta  de  un  Genero  Guayanes 

Carmen  UUoa  Ulloa 

Missouri  Botanical  Garden,  l\0.  Box  299,  St.  Louis,  Missouri  63166-0299,  U.S.A. 


David  A.  Neill 


.  U.S.A., 


Ulloa  Ulloa  & 


A  New  Species  of  Pedicularis  (Scrophulariaceae)  from 
the  Yaoshan  Mountain,  Yunnan,  China 


EH3  5LR, 


Volume  16,  Number  2 
2006 


Volume  16,  Number  2,  pp.  161-202  of  NOVON  was  published  on  20  July  2006. 


www.  mbgpr  ess .  org 


CONTENTS 


Missouri 


Botanical 

Garden 


A  Journal  for  Botanical  Nomenclature 


Volume  16,  Number  3 
October  2006 


Novon,  A  Journal  for  Botanical 
Nomenclature  from  the  Missouri 
Botanical  Garden 


The  mission  of  the  Missouri  Botanical  Garden  is  to  discover  and  share  knowledge 
about  plants  and  their  environment  in  order  to  preserve  and  enrich  life, 


Novon  publishes  short  articles  whose  primary  purpose  is  the  establishment  of  nomencla¬ 
ture  in  vascular  plants  and  bryophytes.  All  articles  are  peer-reviewed  by  qualified,  indepen¬ 
dent  reviewers. 

Manuscripts  must  fully  state  and  justify  the  reasons  for  proposing  nova.  These  may  include 
detailed  comparisons  with  similar  taxa,  short  keys  to  similar  taxa,  illustrations  to  similar  taxa, 
and  mechanical  nomenclature  reasons,  among  others.  Manuscripts  whose  primary  purpose  is 
other  than  establishment  of  new  nomenclature,  which  usually  are  longer  manuscripts,  cannot 
be  accepted  for  review.  These  include  reviews,  revisions,  monographs,  or  other  papers  that 
incidentally  include  nova. 

Manuscripts  must  follow  the  guidelines  in  the  Checklist  for  Authors  and  be  accompanied  by 
a  completed  copy  of  the  Checklist,  The  Checklist  may  be  downloaded  from  the  Garden’s  web 
site,  www.mbgpress.org,  or  authors  may  contact  the  managing  editor  to  request  a  copy. 

Novon  will  not  knowingly  accept  manuscripts  that  have  been  simultaneously  submitted  to 
other  journals  for  consideration  or  previously  published  in  some  form  elsewhere. 


Victoria  C.  Hollowell 

Editor 

Beth  Parada 

Managing  Editor 

Diana  Gunter 

Associate  Editor 

Ihsan  A.  Al-Shehbaz 

Consulting  Editor 

Roy  E.  Gereau 

Consulting  Editor 

John  Pruski 

Consulting  Editor 

Carmen  Uiloa  Ulloa 

Consulting  Editor 

George  Yatskievych 

Consulting  Editor 

Kanchi  N.  Gandhi 

Nomenclature  Consultant 

Nicholas  J.  Turland 

Nomenclature  Consultant 

Barbara  Mack 

Editorial  Assistant 

Monica  Anderson 

MBG  Press  Assistant 

Novon  is  included  in  the  subscription,  price  of  the 
Annals  of  the  Missouri  Botanical  Garden.  Price  for 
2006:  $160  per  year  U.S.A.;  $170  Canada  and 
Mexico;  $195  all  other  countries.  Four  issues  per 
volume. 

Postal  address:  Subscriptions:  ANNALS  OF  THE 
Missouri  Botanical  Garden,  P.0.  Box  1897,  Law¬ 
rence,  KS  66044-8897;  Manuscript  submissions: 
Novon,  Missouri  Botanical  Garden,  RQ,  Box  299, 
St.  Louis,  MO  63166-0299. 


©  Missouri  Botanical  Garden  2006 


quirements  of  ANSI/NISO  Z39.48-1992. 

Novon  (ISSN  1055-3177)  is  published  quarterly 
by  the  Missouri  Botanical  Garden,  2345  Tower 
Grove  Avenue,  St  Louis,  MO  63110.  Periodicals 
postage  paid  at  St  Louis,  MO  and  additional  mail¬ 
ing  offices  (USJPS  #006-777). 

POSTMASTER:  Please  send  address  changes  to 
Annals  Of  the  Missouri  Botanical  Garden,  P.0.  Box 
1897,  Lawrence,  KS  66044-8897. 

Information  on  the  contents  of  Novon  is  present¬ 
ed  in  SciSearch®,  Research  Alert®,  Current  Con¬ 
tents'/Agriculture,  Biology,  and  Environmental 
Sciences,  APT  Online,  the  CAB  Abstract/Global 
Health  database,  and  the  ISI®  database. 


Typesetting  by  Allen  Press,  Inc.  using  Advent3B2  automatic  paging  system. 


Volume  16  NOVON 
Number  3 
2006 


Nomenclatural  Innovations  in  Neotropical  Salicaceae 

Mac  H.  Alford 


Neotropical  Salicaceae 


Astragalus  asotinensis  (Fabaceae),  a  Newly  Discovered 
1  Idaho,  United  States 


2006 


3 


mui 


fiillr 


(Fig.  1). 


y?t;[PS 


314 


A  New  Species  of  Cytinus  (Cytinaceae)  from  South  Africa  and 
Swaziland,  with  a  Key  to  the  Southern  African  Species 


^Cytinusftom  Southern  Africa 


Una  Especie  Nueva  de  Thompsonella  (Crassulaceae)  del  Estado  de 
Michoacan,  Mexico 


Pablo  Carrillo-Reyes 


Wigandia  ecuadorensis  (Hydrophyllaceae),  una  Nueva  Especie  del 
Bosque  Muy  Seco  Tropical  al  Occidente  de  Ecuador 


Meliosma  stellata ,  una  Nueva  Especie  de  Sabiaceae  de  Ecuador 


A  New  Combination  for  Thrasya  robusta  in  Paspalum 
(Poaceae,  Paniceae) 


16:  331. 


Thoreauea  guerrerensis  (Apocynaceae,  Apocynoideae), 
una  Nueva  Especie  de  Guerrero,  Mexico 


Three  New  Mesoamerican  Taxa  of  Dichaea  with  Caducous  L 

Robert  L.  Dressier 


?  mm,  claw  5.5-6  X  3.5-  luol  and 


lip,  ligule  1.5-2  X  1.5-2  mm,  s 


Five  New  Subspecies  of  Braya  (E 


of  B. 


/ 


350 


The  Genus  Callitriche  (Callitrichaceae)  in  Asia 


,  Glos,  GL5  4SQ,  United  ! 


New  Combinations  in  Actinidiaceae  from  China 


Li  Xin-Wei 

Herbarium  (HIB),  Wuhan  Botanical  Garden  of  Chinese  Academy  of  Sciences,  Wuhan  430074 
People’s  Republic  of  China;  Graduate  School  of  the  Chinese  Academy  of  Sciences, 


College  of  Pharmacy,  University  of  Illinois  at  Chicago,  833  S.  Wood  Street, 
Chicago,  Illinois  60613.  I’.S.A. 


Li  et  al. 


Two  New  Species  of  Carex  sect.  Racemosae  (Cyperaceae)  from  China 


Calciphila ,  a  New  Genus  in  African  Asclepiadeae  (Apocynaceae, 
Asclepiadoideae),  and  Taxonomic  Rectifications  in  Cynanchum 


of  Calciphila  gillettii. 


T\  I'K: 


371 


Begonia  hahiepiana ,  a  New  Species  of  Begonia  Section  Sphenanthera 
(Begoniaceae)  from  Vietnam 


Mark  C.  Tebbitt 

Brooklyn  Botanic  Garden,  1000  Washington  Avenue,  Brooklyn,  New  York  11225,  U.S.A. 


374-377.  Pi 


376 


Novon 


377 


Isostigma  sparsifolium  (Asteraceae,  Heliantheae,  Coreopsidinae), 
a  New  Species  from  Goias,  Brazil 


I  i  ■*  ,y 


Crataegus  spes-aestatum ,  a  New  Species  in  Series  Punctatae  (Rosaceae), 
and  Six  New  Varietal  Names  from  the  Missouri  Crataegus  Flora 


,  The  University  of  We 


Species  of  Scutellaria  (L 


3)  from 


Scutellaria  from  Mesoamerica 


393 


No  von 


402 


Novon 


mil 


A  New  Species  and  Conservation  Status  of  Schindleria  (Phytolaccaceae) 
from  Peru 


(TEX), 


A  New  Species  of  Ludia  (Salicaceae)  from  Madagascar’s  Eastern 
Littoral  Forest 


P.O.  Box  299,  Si.  I 


Novon 


Mimosoideae)  from  Central  and  North  America 


David  S.  Seigler 


Novon  16:  413-420.  Published  on  7  November  2006. 


iijiJ 


414 


417 


BM,  F). 


418 


et  al. 


A  New  Species  of  Phyllanthus  (Phyllanthaceae)  from 


Northeastern  Brazil 


Species  of  Ferns  from  the  Rio  Cenepa  Area, 


Illlllilllllll 


The  New  Name  Johnstonalia  Substitutes  for  Johnstonia  (Rhamnaceae) 


5fS£ 


A  New  Name  in  Chinese  Bolbitidaceae 


Xing  Fuwu 


ina 


A  New  Lectotypification  for  Magnolia  fistulosa  (Magnoliaceae) 


Guangzhou  510650,  People’s  Republic  of  China. 


(Lauraceae):  A  New  Species  from 


Guangdong,  China 


441 


Impatiens  X  pacifica  (Balsaminaceae),  a  New  Hybrid  Jewelweed  from 
the  Pacific  Northwest  Coast  of  North  America 


Novon 


447 


www.mbgpress.  org 


CONTENTS 


■^[k  Missouri 

I— ^  /^4-n^ i  ri  nl 


Botanical 

Garden 


A  Journal  for  Botanical  Nomenclature 


NUMBER 


2006 


Volume  16,  Number  4 
December  2006 


Novon,  A  Journal  for  Botanical 
Nomenclature  from  the  Missouri 
Botanical  Garden 


The  mission  of  the  Missouri  Botanical  Garden  is  to  discover  and  share  knowledge 
about  plants  and  their  environment  In  order  to  preserve  and  enrich  life. 


Novon  publishes  short  articles  whose  primary  purpose  is  the  establishment  of  nomencla¬ 
ture  in  vascular  plants  and  bryophytes.  All  articles  are  peer-reviewed  by  qualified,  indepen¬ 
dent  reviewers. 

Manuscripts  must  fully  state  and  justify  the  reasons  for  proposing  nova.  These  may  include 
detailed  comparisons  with  similar  taxa,  short  keys  to  similar  taxa,  illustrations  to  similar  taxa, 
and  mechanical  nomenclature  reasons,  among  others.  Manuscripts  whose  primary  purpose  is 
other  than  establishment  of  new  nomenclature,  which  usually  are  longer  manuscripts,  cannot 
be  accepted  for  review.  These  include  reviews,  revisions,  monographs,  or  other  papers  that 
incidentally  include  nova. 

Manuscripts  must  follow  the  guidelines  in  the  Checklist  for  Authors  and  be  accompanied  by 
a  completed  copy  of  the  Checklist.  The  Checklist  may  be  downloaded  from  the  Garden’s  web 
site,  www.mbgpress.org,  or  authors  may  contact  the  managing  editor  to  request  a  copy, 

Novon  will  not  knowingly  accept  manuscripts  that  have  been  simultaneously  submitted  to 
other  journals  for  consideration  or  previously  published  in  some  form  elsewhere. 


Victoria  C.  Holltawell 

Editor 

Beth  Parada 

Managing  Editor 

Diana  Gunter 

Associate  Editor 

Ihsan  A.  Al-Shehbaz 

Consulting  Editor 

Roy  E.  Gereau 

Consulting  Editor 

John  Pruski 

Consulting  Editor 

Carmen  Ulloa  Ulloa 

Consulting  Editor 

George  Yatskievych 

Consulting  Editor 

Kanchi  N.  Gandhi 

Nomenclature  Consultant 

Nicholas  J.  Turland 

Nomenclature  Consultant 

Barbara  Mack 

Editorial  Assistant 

Monica  Anderson 

MBG  Press  Assistant 

Novon  is  included  in  the  subscription  price  of  the 
Annals  of  the  Missouri  Botanical  Garden.  Price  for 
2007:  $165  per  year  U.S.A.;  $175  Canada  and 
Mexico;  $200  all  other  countries.  Four  issues  per 
volume. 

Postal  address:  Subscriptions:  ANNALS  of  THE 
Missouri  Botanical  Garden,  P.Q..  Box  1897,  Law¬ 
rence,  KS  66044-8897;  Manuscript  submissions: 
Novon,  Missouri  Botanical  Garden,  P.01.  Box  299', 
Si  Louis,  MO  63166-0299. 

Electronic  addresses: 

Editorial  queries;  novon@mobot.oig 
Orders  for  back  issues:  mbgpress@mob0t.org 
Web  site:  http ://www,mbgpreiSS •  oig 

©  Missouri  Botanical  Garden  2006 
Typesetting  by  AUen  Press,  Inc.  using  Advent3B2  a 


©  Novon  is  printed  on  paper  that  meets  the  re¬ 
quirements  of  ANSVNISO  Z39.48-1992. 

Novon  (ISSN  1055-3177)  is  published  quarterly 
by  the  Missouri  Botanical  Garden,  2345  Tower 
Grove  Avenue,  St.  Louis,  MO  63110.  Periodicals 
postage  paid  at  St,  Louis,  MO  and  additional  mail¬ 
ing  offices  (U5PS  #006-777), 

POSTMASTER:  Please  send  address  changes  to 
Annals  of  the  Missouri  Botanical  Garden,  P.O.  Box 
1897,  Lawrence,  KS  66044-8897. 

Information  on  the  contents  of  Novon  is  present¬ 
ed  in  SciSearch®,  Research  Alert®,  Current  Com- 
tents®/Agriculture,  Biology,  and  Environmental 
Sciences,  APT  Online,  the  CAB  Abstract/Global 
Health  database,  and  the  ISI®  database. 


Volume  16  NOVON  ^||L* 
Number  4 
2006 


Una  Nueva  Variedad  de  Lophostachys  guatemalensis  (Acanthaceae)  de 
Interesante  Distribucion  Geografica 


A  New  Species  of  Brachystelma  (Apoc 
Tropical  Africa 


e)  from  South 


/  of  Cape  Town,  7701 


Volume  16,  Number  ■ 


Three  New  Species  of  Euphorbia  (Euphorbiaceae)  from  South 
Tropical  Africa 


KJJUUl, 


Five  New  Species  of  Guarea  (Meliaceae),  Two  from  the  Guarea  glabra 
Vahl  Complex,  in  Central  America 


New  Combinations  in  Melaleuca  for  Australian  Species  of 


,  GPO  Box  1600,  ( 


472 


474 


Senecio  glandulifer  (Senecioneae,  Asteraceae):  Una  Nueva  Especie  del 
Nordeste  de  Argentina 


i  (UNNE-CONICET),  I 


Noyon  16:  476-479.  Published 


19  Dec 


de  2. 3-2. 5  mm;  e 


,  de  0.6-0.8  mm.  ] 


478 


mii 


479 


A  New  Species  of  Dendropanax  (Araliaceae)  from  the  State  of 
Espfrito  Santo,  Brazil 


Three  New  Species  of  Elvasia  (Ochnaceae)  from  the  Brazilian  Atlantic 
Forest,  with  an  Emended  Key  for  Subgen  us  Hostmannia 


19  December  2006. 


A  New  Species  of  Baccharis  (Asteraceae,  Astereae)  from 
Santa  Catarina,  Brazil 


Daniel  A.  Giuliano 


491 


“•  Hi 


Taxonomic  Identity  of  Musa  rosea  (Musaceae)  in  Southeast  Asia 


496 


Novon 


A  New  Species  of  Bulbophyllum  (Orchidaceae)  from  Yunnan,  China 


of  Sc 


1  "|N|,|  ;  i ,  '  h  x "  ''!!  , v,  I'1"1/  ' 


Volume  16,  Number  4 


Jin 

Bulbophyllum  from  Yunnan,  China 


Bulbophyllum  Thou.  Dansk  Bot.  Ark.  33(3):  80- 
si,  Z.  H.  1999.  Bulbophyllum.  Pp.  164-257 


iiiiii 


Novitates  Gabonensis  59:  Combretum  wilksii  (Combretaceae),  a  New 
Species  from  Gabon 


Card  C.  H.  Jongkind 


Myrcia  inconspicua  (Myrtaceae),  a  New  Species  from 
Espirito  Santo,  Brazil 


31270-110,  Belo  Horizonte,  Minas  Gerais,  Brazil,  sobral@ufmg.br 


Croton 


(Section  Eluteria :  Euphorbiaceae),  a  New  Species 
to  the  Yucatan  Pe 


A  New  Subspecies  of  Solanum  scabrum  Miller  Found  in  Uganda 

Eunice  Apio  Olet 


!  Kdre  A.  Lye 

lagement,  Box  5003,  NO-1432  Aas,  ] 


fr  ni  in  r| 


A  New  Central  African  Species,  CX 
from  the  Sources  of  t 


:  35,  B-9000  Gent,  I 


Volume  16,  Number  4 


Reynders  et  al. 

Cyperus  vandervekenii  from  Rwanda 


513 


Kyllinga  benir 


Four  New  Myrtaceae  from  Eastern  Brazil 


Marcos  Sobral 


.  Carlos  6627,  31270-110,  BH<>  I 


525 


Novon  16:  330-532.  Pu 


Two  New  Species  of  Bauhinia  (Fabaceae,  Caesalpinioideae,  Cercideae) 
from  Southeastern  Mexico  and  Mesoamerica 


,  04510,  Mexico  D.F. 


0.7-1. 4 


Volume  16,  Number  I.  |>|>.  1-160  of  NOVON  was  published  on  25  May  2006. 

\  olume  16,  Number  2,  pp.  161-292  of  N<>\  ON  was  published  on  26  July  2006. 
Volume  16,  Number  3,  pp.  293^148  of  NOVON  was  published  on  07  November  2006. 
Volume  16,  Number  4,  pp.  449-538  of  NOVON  was  published  on  19  December  2006. 


www.  mbgpr  ess .  or  g 


CONTENTS